They are diurnal and arboreal, and forage at a height of from 39 to 66 feet. They search for insects in epiphytic bromeliads, in the leaf litter of vine tangles, on bark, and in tree holes and crevices. They do not normally come to the ground. Nectar is an important part of their diet from August to November. They prefer primary tropical forest, but have been found in secondary forest and areas under partial cultivation. They like areas in trees where interlacing branches, vines and epiphytes provide optimum shelter and an abundance of insects and small invertebrate prey. They sleep at night in tree holes, or occasionally in vines or epiphytes. They leap from branch to branch with great agility. Groups occupy home ranges of up to 200 hectares. A portion of the home range appears to be a territory that is defended from other groups. Groups consist of from 2 to 11, usually 3 to 7, related individuals, but temporary associations of 15 or 16 have been observed. The basic group is thought to consist of a mated pair plus their young of one or more years. Although adults of the same sex have been reported to be extremely aggressive toward one another, even fighting to the death if kept together, numerous observations now indicate that natural groups frequently contain more than one adult of each sex. The dominant adult male and female of a group form a permanent pair bond, are equal in rank, and share responsibility for raising the young.